B.A. Biology, Gustavus Adolphus College, St. Peter, Minnesota, 1995
M.S. Biology/Parasitology, South Dakota State University, Brookings, South Dakota, 1998
D.V.M., University of Minnesota, College of Veterinary Medicine, St. Paul, MN, 2001
Ph.D., University of Minnesota, College of Veterinary Medicine, St. Paul, MN, 2007